Melolonthinae (subfamily) (Cockchafer)
- tarsal claws equal and fixed
- mandibles completely concealed from above
- only one pair of abdominal spiracles (no. 8) visible beneath the edges of the elytra
- head and pronotum simple (lacking horns or other protrusions)
- tarsal claws often toothed
- the externally visible sclerotised labrum
